Fillion has worked in several theatre, television and film productions, including the improvised soap opera "Die Nasty".
Fillion was Joey Buchanan in "One Life to Live" and was nominated in 1996 for a Daytime Emmy Award in the "Outstanding Younger Actor" category. He was in the sitcom "Two Guys, a Girl, and a
Pizza Place". He was also cast as "the Minnesota Ryan" in Steven Spielberg's "Saving Private Ryan".
Fillion starred as Captain Malcolm Reynolds in the Joss Whedon science fiction television series Firefly in 2002 and he won the "Cinescape Genre Face of the Future Award - Male" award by the Academy of Science Fiction, Fantasy & Horror Films, USA.
Fillion had a role in the film "Serenity" in 2005. He had a recurring role as Caleb in the final season of Whedon's series Buffy the Vampire Slayer.
Fillion has lent his voice to the animated series King of the Hill in 2001, the video game Jade Empire, and the animated series Justice League Unlimited in 2005 and 2006. Fillion is currently filming "White Noise 2: The Light".
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